Kazuma is a native Alter user who makes his living as a mercenary so that he and his close friend, Kanami Yuta, can live together. Kazuma's friend Kunihiko Kimishima gives him a mission to defeat another native Alter user. However, a man known as Ryuho from HOLY captures Kazuma in the aftermath. Ryuho tries to interrogate Kazuma over an unknown Alter user (later dubbed Crystal) who killed his mother and dog. Using a woman from HOLY named Mimori Kiryu as a hostage, Kazuma manages to return to his house. When native Alter users are captured by HOLY, Kazuma enters a forest where he finds Crystal. After fighting Crystal, Kazuma's Alter receives a power up. Shortly afterwards, it is revealed the Great Uprising was caused by a similar event in a parallel universe where all creatures are independent thinking Alters. Kazuma goes on a rampage against HOLY after Kinishima dies while helping him during one of his fights. Shortly afterwards, Kazuma and Ryuho fight, but during the battle, they open a gateway to the parallel universe, causing another uprising and attracting the attention of the Mainland government.
Both Kazuma and Ryuho are thought to be dead after the fight. While Kazuma hides in underground fights, Ryuho suffers amnesia. With help from an old friend, HOLY member Straight Cougar, Kazuma recovers his will to protect Kanami who is searching for him. Ryuho also recovers his memories while fighting HOLY users attacking natives and ends up becoming one of Kazuma's allies. The Mainland sends its agent, Kyouji Mujo, to the Lost Ground in an attempt to exploit the power of the parallel universe for economic gain. Mujo develops delusions of grandeur after becoming the de facto head of the Lost Ground government and absorbing the crystal entity from 'the other side' to add to his already formidable Alter power. Both he and the Crystal entity are defeated by the separate efforts of Kazuma and Ryuho respectively, as both increase their Alter powers by taking energy from the parallel world. The Mainland once again attempts to exert control over the Lost Ground by sending in a fleet of refined Alter users under their control. However, they are defeated by Kazuma and Ryuho.
After defeating the Mainland forces, Kazuma and Ryuho engage in one final battle to settle their rivalry. As one of them is about to declare victory, the screen fades to white. In an epilogue, taking place years later, they remain the protectors of the Lost Ground, defending it from Mainland invaders, while an older Kanami awaits their return.
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